Publications (updated Nov. 2016)




· Mlynarek, J.J., J.-H. Kim, and S.B. Heard. 2016. Identification of leaf mining insects via DNA recovered from empty mines. FACETS, in press.  (preprint)

· Pureswaran, D., R. Johns, S.B. Heard, and D. Quiring. 2016. Paradigms in eastern spruce budworm (Lepidoptera: Tortricidae) population ecology: A century of debate. Environmental Entomology, online early, DOI:

· Shibel, Z. and S.B. Heard. 2016. Synergistic and additive effects of drought stress and simulated herbivory on two goldenrods, Solidago altissima L. and S. gigantea Ait.  Botany 94: 635-642.  (pdf)

· Dearborn, K.W., S.B. Heard, J. Sweeney, and D.S. Pureswaran. 2016. Displacement of Tetropium cinnamopterum (Kirby) (Coleoptera: Cerambycidae) by its invasive congener, the brown spruce longhorn beetle, Tetropium fuscum (Fabricius).  Environmental Entomology 45:848-854. (pdf)

· Rhainds, M., S.B. Heard, C. Hughes, W. MacKinnon, K. Porter, J. Sweeney, P. Silk, I. DeMerchant, S. McLean, and G. Brodersen. 2015. Evidence for mate-encounter Allee effect in an invasive longhorn beetle (Coleoptera: Cerambycidae). Ecol. Entomol. 40:829-832. (pdf)

· Grégoire, D.M., D.T. Quiring, L. Royer, S.B. Heard, and É. Bauce. 2015. Indirect host-mediated effects of an exotic phloem-sap feeder on a native defoliator of balsam fir.  For. Ecol. Manag. 341:1-7. (pdf)

· Rhainds, M. and S.B. Heard. 2015. Sampling procedures and adult sex ratios in spruce budworm. Entomol. Exper. Applic. 154:91-101. (pdf)

· Heard, S.B. 2014. On whimsy, jokes, and beauty: can scientific writing be enjoyed?  Ideas Ecol. Evol. 7:64-72. (pdf)

· Oke, O.A., S.B. Heard, and J.T. Lundholm. 2014. Integrating phylogenetic community structure with species distribution models: an example with plants of rock barrens.  Ecography 37:614-625 online early (pdf)

· Carleton, R.D., S.B. Heard, and P.J. Silk. 2013. A simulation approach to assessing sampling strategies for insect pests: an example with the balsam gall midge.  PLOS One 8(12):e82618. (pdf)

· Carleton, R.D., P.J. Silk, E.S. Eveleigh, S.B. Heard, and C. Dickie. 2013. A sampling unit for estimating gall densities of Paradiplosis tumifex (Diptera: Cecidomyiidae) in Christmas tree stands. Can. Entomol. 145:343-349. (pdf)

· Heard, S.B. and E.K. Kitts. 2012. Impact of attack by Gnorimoschema gallmakers on their ancestral and novel Solidago hosts.  Evol. Ecol. 26:879-892 (pdf).

· Woods, E.C., A.P. Hastings, N.E. Turley, S.B. Heard, and A.A. Agrawal. 2012. Adaptive geographical clines in the growth and defense of a native plant. Ecol. Monogr. 82:149-168 (pdf).

· Heard, S.B. 2012. Use of host-plant trait space by phytophagous insects during host-associated differentiation: the gape-and-pinch model. Int. J. Ecol. 2012:ID192345 (pdf).

· Holmes, S.E., E.G. Kettela, P.C. Nigam, S.B. Heard, and R.C. Johns. 2012. Temporal variations in abiotic conditions exert little influence on diurnal activity of late-instar spruce budworm larvae on balsam fir.  Can. Entomol. 144:396-405 (pdf).

· Ancheta, J. and Heard, S.B.. 2011. Impacts of insect herbivores on rare plant populations.  Biol. Conserv. 144:2395-2402 (pdf).

· Heard, S.B. and J. Ancheta. 2011. Effects of salinity and temperature on ex situ germination of the threatened Gulf of St. Lawrence Aster, Symphyotrichum laurentianum Fernald (Nesom). Plant Species Biol. 26:158-162 (pdf).

· Rhainds, M.,  S.B. Heard, J. Sweeney, P. Silk and L. Flaherty. 2010.  Phenology and spatial distribution of native and exotic Tetropium longhorned beetles (Coleoptera: Cerambycidae).  Environ. Entomol. 39:1794-1800 (pdf).

· Carleton, D., D. Quiring, S.B. Heard, C. Hébert, J. Delisle, R. Berethiaume, E. Bauce, and L. Royer. 2010.  Density-dependent and density-independent responses of three species of Telenomus (Hymenoptera: Scelionidae) parasitoids of hemlock looper eggs.  Entomol. Exper. Appl. 137:296-303 (pdf).

· Ancheta, J., S.B. Heard, and J.W. Lyons. 2010.  Impacts of salinity and simulated herbivory on survival and reproduction of the threatened Gulf of St. Lawrence Aster, Symphyotrichum laurentianum. Botany 88: 737-744 (pdf).

· Hendrix, S.D., K.W. Kwaiser, and S.B. Heard. 2010.  Bee communities (Hymenoptera: Apoidea) of small Iowa hill prairies are as diverse and rich as those of large prairie preserves.  Biodivers. Conserv. 19:1699-1709 (pdf).






The Heard Lab

Understanding ecological controls on the evolution of biodiversity

Plant images © Geoff Galicio, Pethan, Randy Nonenmacher, Elke Love